I think that right now you should run the most current OS that is stable (OS 10.6.x). OS 10.7 will probably run very well on your new machine after the kinks are worked out. 10.8 will probably come out 1 to 2 years after 10.7 is stable. This cycle for OS X has been going on for a decade.
Each new OS will provide new features and probably speed improvements. They will also bring the usual upgrade requirements for drivers for existing hardware, and compatibility issues with existing programs. These updates will take whatever time the 3rd party vendors choose to take. Some programs or drivers may never come out or get updated for a new OS and you'll either have to find substitues, or keep a copy of the old OS on a bootable drive, or just decide not to upgrade at all.
Run the best stable OS you can for now, and worry about what's going to happen in a couple of years for then.
Except for the lengthy time that WinXP was around (far too long as the work/world was changing), most OS's are upgraded every couple of years. I hear Windows 8 is in development, and Ubuntu Linux 11 is coming out on the 28th of this month. It's not supposed to end.