you can install Lion
I would look for Mac Pro 4,1 2009.
For me, I added 3 x SSDs. I put two on Sonnet Tempo (you can get the Pro but 2x $)
I then boot from SSD #1 in drive bay 0.
I put two 256GB SSD on Sonnet Tempo.
I don't use LR5 but I would put its catalogue on one SSD
I would host photo images and imports on WD 2TB in drive bay #3 (bay #2 has other user files and media)
US$ for following:
120GB SSD Samsung 840 for system $96
240GB of same $170
500GB $320
Sonnet Tempo $150 / Pro version is for RAID performance 900MB/sec $300
ATI 5770 $250
8 x 2GB of RAM if you have less than that, more is better, $30 2 x 2GB
That will with those SSDs run smoothly. Can't hurt to do 5355s as they are better and for $100.
2x2GB FBDIMM DDR2 667MHz @ $30
http://www.amazon.com/BUFFERED-PC2-5300-FB-DIMM-APPLE-Memory/dp/B002ORUUAC/
Mac Pro 8GB Kit Komputerbay 8GB (2x 4GB) DDR2 667MHz ECC FB-DIMM (240 PIN)
http://www.amazon.com/Komputerbay-PC2-5300F-Buffered-FB-DIMM-Heatspreaders/dp/B0 05HIWD5U/
eBay (Europe)
http://viewitem.eim.ebay.dk/KOMPUTERBAY-8GB-2X4GB-MEMORY-for-APPLE-MAC-PRO-EARLY -2008-31-DDR2-800MHz/140962093586/item?transId=808969265004
A mini does not cut it for me. I could get along with an iMac but I'd wait for newer model at this point.
RAM $120
SSDs $400
Sonnet Tempo $150-300
5355 $100
ATI 5770 $250
That is about what a 2009 can be had for though I did it one item per month over 6 months